A passage of dark, wet rock leading to a floor of soft orange mud, lit from one side and fading into darkness, an impression of the deep, hard-to-reach chambers of a South African cave system like the one where Homo naledi was found.

Human RelativesPleistocene

Homo naledi

A small-brained human relative known from a single deep cave, and the centre of an argument about burial

About 335 000 to 236 000 years ago Reconstruction, not a photograph
Lived about 335 000 to 236 000 years ago dated in 2017; the burial team now writes 335 000 to 241 000 [1]
Where Rising Star cave system, South Africa not found outside the Cradle of Humankind [14]
Found 13 September 2013 by the cavers Hunter and Tucker [2]
Named 2015 naledi means star in the Sotho language [2]
Fossils over 1 550 specimens at least 15 individuals, plus at least 3 from the Lesedi Chamber [1]
Height about 1.44 m [1]
Weight 39.7 to 55.8 kg range of estimates [1]
Brain 465 to 610 cm³ modern people 1 270 to 1 330 cm³ [2]
Depth of the chamber about 30 m reached today through squeezes as tight as 20 cm [3]
Burial and engravings claimed, and disputed reviewers split on burial; engraving study rated incomplete [5]

About the pictures. Two paintings, neither showing Homo naledi itself: a dark cave passage with a floor of orange mud, after the orange-red muddy sediments of the Rising Star cave system, and scratched crossing lines on pale rock, after the disputed marks on a natural pillar there. Both are impressions, not copies. The real chambers are far narrower and lie in total darkness, so the light and the width of the passage are imagined, and the pattern of the lines is invented. The age and maker of the real marks are unproven. No skin or hair survives, so nothing is known of the colour of its skin or hair.

Homo naledi was a small human relative known from only one place, the Rising Star cave system in South Africa’s Cradle of Humankind, where its bones date to between about 335 000 and 236 000 years ago. They are the largest collection of a single early human species found in Africa, and they pair a small brain with hands and feet much like ours. Its discoverers argue that it buried its dead and engraved cave walls. Other researchers, and some reviewers of those papers, say the evidence does not show that.

Thirty metres down, through a 20 cm gap

Two cavers, Hunter and Tucker, found the bones on 13 September 2013. The Dinaledi Chamber lies about 30 m below the surface and about 80 m in a straight line from the nearest present entrance, and it was always in total darkness. Reaching it today means climbing a jagged fallen block, then descending about 12 m through a fissure with squeezes as tight as 20 cm. Berger’s team recovered over 1 550 specimens from at least 15 individuals, and at least three more came from the nearby Lesedi Chamber. The species was named in 2015, from the Sotho word for star.

Its small brain suggested an age of 1 to 2 million years. Dating in 2017 gave a far younger answer, so it lived at the same time as larger-brained humans.

An old body plan, late in time

Adults stood about 1.44 m and weighed about 40 to 56 kg. The brain measured 465 to 610 cm³, less than half that of modern people. The shoulders and pelvis resemble those of Australopithecus, with shoulder blades set high and short collarbones, while the hands and feet are like those of Homo. The thumb was strong, built for a precise grip, yet the finger bones are curved, which is read as a sign of climbing. The legs suited long-distance walking with a human-like stride but not endurance running. No tools are firmly linked to the bones, little is known of its diet and habitat, and its place in the family tree is unresolved.

Did it bury its dead?

The idea goes back to 2015, when the team noted that bodies seemed to reach the chamber whole, in a place closed to large predators. In 2023 Berger and colleagues went further in the journal eLife, arguing that bodies were placed in dug pits and covered. They describe two features in which most bones come from a single body, one of them also holding teeth of at least two other individuals, with bones still articulated or at angles they could not hold unsupported, in sediment unlike the surrounding floor. They say slumping, water and carnivores cannot explain the pattern.

eLife publishes papers alongside their reviews instead of accepting or rejecting them. Its 2023 assessment recorded a “strong consensus” among four reviewers that the methods, data and analyses did not support the conclusions. Martinón-Torres and colleagues argued that natural processes had not been ruled out, nor pits shown to be dug and filled. In 2025 Foecke, Queffelec and Pickering reanalysed the sediment chemistry in the 2023 version and found that all sample groups overlapped, with no detectable difference between the supposed graves and the surrounding sediment.

The team revised the paper, reworking the sediment chemistry in answer to such criticism, and now reports trace-element signatures found only with the bones. The final version appeared in September 2025. One reviewer who had criticised the first version now finds sufficient evidence for repeated burials. Another considers the evidence incomplete, pointing to a 2021 study that suggests an easier way in before the fallen block came down, and hopes that independent teams will analyse the site. eLife’s assessment calls it an improved case and leaves the hypothesis open.

Marks on a pillar, and no male marker

The same team reported at least 46 marks, grooves of about 5 to 15 cm, some crossing, on a natural pillar at the entry to a passage into the Dinaledi Chamber. They have not been dated directly: their age is borrowed from the bones, and Homo naledi is named as the maker because it is the only human relative known from that space. eLife rates the study important but incomplete. The team says the lines differ from the natural weathering of the rock beside them. Garate says weathering or animal claws can make similar marks, and one reviewer concluded that nothing refutes the claims and nothing convincingly confirms them.

A 2026 study led by Madupe extracted proteins from 23 teeth of at least 20 individuals and found no male marker. That could mean only females were placed in the cave, or that an isolated population had lost or altered the gene behind the marker.

Where it sits in deep time

Homo naledi: About 335 000 to 236 000 years ago, in the Pleistocene.

Homo naledi

Earth forms, 4.57 billion years agoToday

Each step to the right is ten times closer to today, so the last few million years get as much room as the first billion.

If Earth's history were one day

Squeeze all of Earth's history, about 4.6 billion years, into 24 hours, starting at midnight. Homo naledi runs from 23:59:53.7 to 23:59:55.5, ending 4.5 seconds before midnight.

On this clock one second is about 52 862 years. All of written history fits into the last tenth of a second.

How tall

Estimated height of Homo naledi next to a modern person.

Reconstructions

A passage of dark, wet rock leading to a floor of soft orange mud, lit from one side and fading into darkness, an impression of the deep, hard-to-reach chambers of a South African cave system like the one where Homo naledi was found. AI-generated painting.
A patch of scratched, crossing lines on a pale rock surface deep inside a cave, lit from one side, an impression of the disputed engravings in the Rising Star cave system attributed to Homo naledi. AI-generated painting.

Questions people ask

Did Homo naledi bury its dead?

That is disputed. Its discoverers argue that bodies were placed in dug pits and covered, which would be the oldest known burials. Other researchers say natural processes have not been ruled out, and one team found that the sediment data in the 2023 version of the paper could not separate the supposed graves from the surrounding floor; the discoverers have since reworked that analysis. Of the reviewers of the final paper in 2025, one accepted the burials and another found the evidence incomplete.

Did Homo naledi make rock engravings?

That is claimed, not established. Its discoverers say that crossed grooves on a pillar in the cave are engravings and that Homo naledi is the likeliest maker, as the only human relative known from that space. But the marks have not been dated directly, and critics say weathering or animal claws can leave similar marks. eLife rates the study as incomplete.

When did Homo naledi live?

Between about 335 000 and 236 000 years ago, according to dating of teeth, flowstone and sediments published in 2017. Earlier guesses of 1 to 2 million years, based on its small brain, were far too old.

How big was Homo naledi?

About 1.44 m tall, with weight estimates from 39.7 to 55.8 kg. Its brain measured 465 to 610 cm³, less than half that of modern people.

Were all the Homo naledi in the cave female?

Possibly. A 2026 study of proteins from 23 teeth of at least 20 individuals found no male marker. That could mean an all-female deposit, or an isolated population in which the gene behind the marker had been lost or altered.

How was Homo naledi discovered?

The cavers Hunter and Tucker found the bones in the Rising Star cave system on 13 September 2013. Berger then organised an excavation, and the species was named in 2015.

Sources

  1. Smithsonian Human Origins: Homo naledi
  2. Wikipedia: Homo naledi
  3. Dirks et al. (2015), Geological and taphonomic context for the new hominin species Homo naledi from the Dinaledi Chamber, South Africa, eLife 4, e09561
  4. Dirks et al. (2017), The age of Homo naledi and associated sediments in the Rising Star Cave, South Africa, eLife 6, e24231
  5. Berger et al. (2025), Evidence for deliberate burial of the dead by Homo naledi, eLife 12, RP89106
  6. eLife (2023), first assessment of Berger et al., Evidence for deliberate burial of the dead by Homo naledi
  7. eLife (2025), peer reviews of Berger et al., Evidence for deliberate burial of the dead by Homo naledi
  8. Berger et al. (2025), An initial report of circa 241,000- to 335,000-year-old rock engravings and their relation to Homo naledi in the Rising Star cave system, South Africa, eLife 12, RP89102
  9. eLife (2025), peer reviews of Berger et al., rock engravings in the Rising Star cave system
  10. Phys.org (2025), New Homo naledi evidence supports intentional burial practices
  11. Martinón-Torres et al. (2024), No scientific evidence that Homo naledi buried their dead and produced rock art, Journal of Human Evolution 195, 103464
  12. Griffith University (2023), No scientific evidence for cognitively advanced behaviours and symbolism by Homo naledi
  13. Foecke, Queffelec & Pickering (2025), No geoarchaeological evidence for deliberate burial by Homo naledi: On best practice for geochemical studies in archaeology and paleoanthropology, PaleoAnthropology 2025(1)
  14. Daily Maverick (2026), Inside the Gauteng caves where rocks reveal the mysteries of Homo naledi
  15. Madupe et al. (2026), Proteomic analysis of dental enamel from 20 Homo naledi individuals shows no male markers, Cell 189, 4415–4424
  16. University of York (2026), Ancient tooth protein reveals 'all-female' fossil site